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- 90-2 form
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A list of research projects funded by CT Sea Grant can be found here.
Currently Funded CT Sea Grant Research (2022-2024)
Project Title: “Copepod Adaptation to Climate Change: No Free Lunch?”
Researchers:
Lead PI: Hans Dam, UConn Dept. of Marine Sciences
Co-PI: Michael Finiguerra, UConn Dept. of Ecology and Evolutionary Biology
Summary: Building on earlier research on the copepod species Acartia tonsa, this research examines the costs of adaptation to future climate conditions that can constrain copepod evolutionary rescue, limit population resilience and consequently reduce fisheries and aquaculture yields. Population fitness is being tested after prolonged exposure to 100 generations of ocean warming, ocean acidification and combined warming and acidification conditions. The research is also testing if adaptation to these climate change conditions diminishes population fitness, and whether there are differences in adaptation costs among ocean warming, ocean acidification and combined warming and acidification conditions. Tests are also being conducted to discover any hidden costs of adaptation to the three climate change conditions manifested as increased susceptibility to toxic phytoplankton and reduced thermal tolerance.
Project Title: “Restoration of Kelp Forests and Associated Ecosystem Services in Long Island Sound”
Researchers:
Lead PI: Sean Grace, Southern Connecticut State University
Co-PI: Colette Feehan, Montclair State University
Summary: The use of “green gravel” as a kelp forest restoration technique is being tested on degraded rocky reefs in Long Island Sound that were formerly dominated by kelp forests but now dominated by algal turfs, to restore the kelp habitat and associated ecosystem services. A partnership with kelp aquaculture farmers is being employed by incorporating locally developed and validated commercial
kelp seed production methods into the project. Local high school students and teachers from the Bridgeport Regional Vocational Aquaculture School and Project Oceanology are also participating in the research as an educational project. The goal of the project is to provide a “proof of concept” for the green gravel technique in Long Island Sound that can be upscaled in the future to restore kelp forests along the degraded Sound coastline. To accomplish broad kelp forest restoration goals, the “green gravel” technique must be piloted locally to establish the conditions under which it will be most effective.
Project Title: “Toward a Deeper Understanding of Human Connections with Ocean Environments: Ocean Identity (OI) as a Novel Construct, Research Instrument, and Assessment Tool”
Researchers:
Lead PI: Miriah Russo Kelly, Southern Connecticut State University
Co-PIs: Jo-Marie Kasinak, Sacred Heart University; Emma McKinley, Cardiff University
Summary: This research seeks to create a model for measuring the construct of ocean identity by providing practical, relevant and timely tools for environmental organization partners. Researchers will first survey academic literature pertaining to the construct of ocean literacy, and then design an instrument to measure it. Data will be collected broadly to validate the instrument and include analysis of differences in race, gender and age. The data will be used for an additional phase of the research that will use Project Limulus as a case study platform for implementing the survey in a way that is appropriate for informal educational settings. The project will also include development of a toolkit that could be used by many organizations to provide informal engagement, outreach and education around ocean issues. In developing a validated, reliable instrument for measuring ocean identity, the researchers seek to establish effective ways for measuring complex human-ocean connections.
Title: “Increasing Sea Bass Abundance in Long Island Sound: A Local or Broader Regional Phenomenon?”
Researchers:
Lead PI: Hannes Baumann, UConn Dept. of Marine Sciences
Co-PIs: Deborah Pacileo, Jacqueline Benway, CT Dept. of Energy and Environmental Protection
Summary: This project is addressing questions raised by the sharply rising abundance of black sea bass in Long Island Sound trawl surveys. The first question is whether the Sound could be a particularly suitable ecosystem for an especially rapid invasion of this warm-water adapted meso-predator. To answer this question, researchers are collecting an analyzing age-structure data for black sea bass from the Sound for length, growth and sex, analyzing historical data and comparing abundance trends from other regions. This project is helping to fill a critical gap in Sound-specific black sea bass age data. This serves the important work of developing accurate updated stock assessments needed by marine fisheries regulators. The relationship of environmental factors such as salinity and temperature to abundance is also being explored, and information on survey efforts in other regions is being gathered and analyzed for indications of abundance trends.
Title: “Risk Averse or Risk Enduring? Understanding the Relationship between Long Island Sound Communities and Flooding to Support Equitable Risk Mitigation Planning”
Researcher:
PI: James Knighton, UConn Dept. of Natural Resources and the Environment
Summary: To advance equitable flooding solutions across Long Island Sound, this research is using socio-environmental models, demographic data and historical records from the National Flood insurance Program (NFIP) to characterize the attitudes and behaviors relevant to a community’s strategy for flood risk mitigation and determine barriers to NFIP participation. Researchers are studying seven Long Island Sound communities protected by coastal surge barriers to determine whether the protection that these barriers provide from moderate floods leads to increased risk when communities relax precautionary behaviors. The project will also develop predictions on how community behaviors might shift in response to future storm conditions of sea level rise combined with more frequent high surge events. This work will be complemented with an examination of the hypothesis that more frequent flooding will drive increased awareness and more community desire to participate in flood protection programs.
Title: “Assessment of PFAS Food Web Uptake in Priority Shellfish Areas: The Roles of Nutrient Status and Trophic Transfer”
Researchers:
Lead PI: Christopher Perkins, UConn Center for Environmental Science and Engineering
Co-PI: Jessica Brandt, UConn Dept. of Natural Resources and the Environment
Summary: This research will address the lack of information about bioaccumulation of per- and polyfluoroalkyl substances (PFAS) in coastal Long Island Sound species, in particular shellfish harvested for human consumption and phytoplankton that form the basis of the Sound’s food web. The study seeks to advance understanding of how nutrient-contaminant interactions mediate PFAS accumulation and trophic transfer to shellfish in coastal food webs by investigating how phytoplankton responses to shifts in nutrient status influence the magnitude of PFAS uptake by these primary producers at the base of the LIS food web, which are direct sources of exposure to CT shellfish species. The research will consist of a baseline assessment of PFAS contaminants in shellfish from regulated beds; an assessment of variations in phytoplankton and shellfish PFAS concentrations; and determination of the relationship between phytoplankton PFAS concentrations and concentrations in blue mussels and wild-caught and farmed shellfish.
Currently Funded LISS Research (2023-2025)
Long Island Sound water quality, salt marsh and public beach characteristics will be examined by marine and social scientists in nine research projects awarded funding by the Connecticut and New York Sea Grant programs (CTSG and NYSG respectively) and the U.S. Environmental Protection Agency’s Long Island Sound Study (LISS) Research Grant Program.
These new projects, which seek information that can be used to improve the conditions of the estuary for humans and wildlife, are being supported by $4.2 million in federal funds. That will be supplemented with matching funds of $2.1 million, for a total research package of more than $6.3 million.
The projects will be conducted over two years beginning this spring. The results will build on the substantial body of research funded through the LISS Research Grant Program administered by CTSG and NYSG since 2008 which has contributed to improved understanding and management of this nationally recognized estuary. Cumulatively, this represents the largest research investment in the Sound, which has been designated an estuary of national significance and one of the most valuable natural resources for both states.
The four CTSG-administered projects are:
- “Testing the effects of vegetation on saltmarsh ecology, services and restoration success: from microbial ecology and biogeochemistry to wildlife conservation,” will be led by UConn professors Christopher Elphick, Beth Lawrence and Ashley Helton, along with Blaire Steven of the CT Agricultural Experiment Station and Min Huang of the CT Department of Energy and Environmental Protection. By creating sediment mounds of varying elevations planted with various species at different densities at Great Meadow Marsh in Stratford, the researchers seek to develop an interdisciplinary understanding of how marsh restoration efforts impact the functioning of these ecosystems and their value for wildlife.
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Rocky Neck State Park in East Lyme is among the most frequently closed or under swim advisories in the state of Connecticut due to high bacteria levels. Judy Benson / Connecticut Sea Grant “Tracking pathogen pathways and fecal bacteria patterns for public beaches suffering with poor water quality grades and closures,” will be led by UConn Professor Michael Whitney with Peter Linderoth of Save the Sound. The researchers will analyze patterns of fecal indicator bacteria in water samples from Green Harbor Beach in New London and Rocky Neck State Park in East Lyme. Because of high bacteria levels, these two public beaches are among the most frequently closed or under swim advisories in the state of Connecticut. The work will include efforts to identify bacteria pathways and public outreach about water quality issues.
- “Assessing temperature mediation of PFAS impacts on coastal fish fitness to inform environmental management,” is being led by professors Maria Rodgers of North Carolina State University and Jessica Brand, Daniel Bolnick, Kat Milligan-McClellan and Milton Levin of UConn. The researchers will examine concentrations of PFAS (poly- and per-fluoroalkyl substances) at different water temperatures in fish populations downstream from the outfalls of public sewage treatment plants. The results will quantify how fish can be expected to respond to exposure to these “forever chemicals” in the Sound over the next five decades.
- “Assessing the impacts of warming and planting strategy on the resilience of restored salt marshes to improve restoration efficacy,” will be led by Sarah Crosby of The Maritime Aquarium at Norwalk, along with Randall Hughes and Nicole Kollars of Northeastern University, Nicole Spiller of Harbor Watch and Earthplace, and LaTina Steele at Sacred Heart University. By enclosing sections of salt marshes within open chambers to increase interior temperatures, the researchers will assess the expected effects of warmer temperatures associated with climate change. The work will include plantings of southern-sourced marsh grass (Spartina) strains to determine impacts on future resilience, and an examination of the genetic mixing of these salt marsh strains to enhance the success of restoration efforts.
The five NYSG-administered projects are:
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Mollusk shells on the shore of Long Island Sound. Dietl’s project will look at the remains of mollusks to understand past ecological conditions in the Sound. Photo: Matthew Pruden “Using geohistorical baselines to assess responses of benthic macroinvertebrate communities to the nitrogen TMDL management intervention in Long Island Sound,” is directed by Gregory Dietl of the Paleontological Research Institution. Scientists will look at the remains of mollusks buried beneath the seafloor to understand past ecological conditions in the Long Island Sound (LIS). The molluscan geohistorical record could provide location-specific information about levels of nitrogen relevant for decades-long time periods that is not available from any other source.
- “Coupled prediction of residential fertilizer use and nitrogen loads to Long Island Sound: An integrated targeting tool for nitrogen-reduction behavior change campaigns,” is led by Robert Johnston of Clark University, along with David Dickson and Jamie Vaudrey of the University of Connecticut, David Newburn of the University of Maryland, Qian Lei-Parent of the University of Connecticut, and Haoluan Wang of the University of Miami. This project will use a survey of households to predict residential fertilizer lawn use for the coastal counties and municipalities across the LIS watershed. A model combining this information with water quality data will be used to inform prospective behavior-change campaigns to identify and prioritize the areas or types of households that would have the greatest impact on reducing nitrogen from lawn fertilizer and its impact on the Sound.
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Team members of the Tzortziou lab collect measurements along Long Island Sound wetlands, near Wheeler Marsh, CT, as well as (Inset) optical measurements using sensor devices. Photo: Tzortziou Bio-Optics Lab “Actionable satellite water-quality data products in LIS for improved management and societal benefits,” is headed by Maria Tzortziou of the City College of New York, along with Joaquim Goes of Columbia University and Melanie Abecassis of the University of Maryland College Park. Human-caused climate change as well as other anthropogenic factors can intensify harmful algal blooms (HABs) in the LIS. Observations of the entire ecosystem, over different seasons and across a range of conditions, including during extreme weather events, can be obtained from satellite data. This work will provide actionable information for water resource management, policy, and decision-making.
- “Evaluating changes in suitable habitat and distribution of cold- and warm-adaptive fish species in a changing Long Island Sound to inform ecosystem-based management,” is led by Yong Chen of Stony Brook University, along with Kurt Gottschall of the Connecticut Department of Energy and Environmental Protection, Kim McKown, and John Maniscalco, at New York State Department of Environmental Conservation. Conditions in the LIS have been shifting due to climate change, affecting water temperature, acidity, oxygen levels, and incidence of HABs. The scientists will evaluate these shifting conditions on the distributional changes of warm-adapted and cold-adapted species of fish in the Sound.
- “Equitable access to Long Island Sound waterfront and beaches through on-demand mobility,” is led by Anil Yazici and Elizabeth Hewitt of Stony Brook University. Some communities on Long Island do not have the mobility means to use and appreciate the LIS waterfront. Project leaders are designing and piloting on-demand shuttles that will facilitate equitable public access to the LIS waterfront. The team will survey users of the shuttle service to identify changes in attitudes toward the LIS environment.
In their words: Long Island Sound Research Projects:
From Sylvain De Guise, director of CTSG: “The continued partnership between Sea Grant programs and EPA will support a nice diversity of innovative and ambitious research projects to benefit both people and ecosystems of Long Island Sound, for mutual benefits—a wise investment, in my opinion.”
From Syma Ebbin, research coordinator for CTSG: “This competition was the largest ever administered, allowing the program to support these diverse, high-quality proposals, all with the capacity to enhance Long Island Sound’s management, health and public benefits.”
From Becky Shuford, director of NYSG: "New York Sea Grant is proud to continue this long-standing partnership with Connecticut Sea Grant and the EPA Long Island Sound Study. This year was the largest research competition to date resulting in the selection of nine excellent and diverse studies that will address priorities related to historical and current water quality conditions, habitat and fisheries health and restoration, and Sound access. The results will have direct benefit to the communities, coasts, critters, and waters of the Long Island Sound Estuary.”
From Lane Smith, research coordinator for NYSG: “This cohort of new research will build on the growing legacy of impactful research that benefits the Long Island Sound and its coastal communities. This continues the fruitful partnership between Sea Grant and the EPA Long Island Sound Study that benefits the LIS ecosystem.”
From David W. Cash, EPA New England regional administrator: “The Long Island Sound estuary is an essential ecosystem that supports communities, economies, and habitats across the region. I’m pleased to say these diverse and innovative Sea Grant projects include a focus on improving the Sound’s water quality, mitigating the effects of climate change, and helping local communities receive more equitable access to the Sound.”
From Lisa F. Garcia, EPA Region 2 regional administrator: “The Long Island Sound is in the center of one of the most densely populated coastlines in the country. This investment will help Long Island Sound communities combat sources of pollution that lead to closing public beaches or contaminating local fish. It will also help communities improve efforts to restore wetland habitat and increase resiliency to climate change by understanding the effects of sea level rise and warming temperatures on valuable marsh habitats. This funding will advance ecological research and play a critical role in improving water quality and reducing pollution, providing lasting results for the wildlife and wetlands in the Sound for years to come.”
Previously Funded LISS Research (2023-2025)
Eight research projects that will examine various facets of the water chemistry and habitat quality of Long Island Sound and potentially yield more effective management decisions have been awarded more than $2.8 million in federal funding through the Long Island Sound Study Research Grant Program.
The projects, supported by a partnership of the Sea Grant programs of Connecticut and New York (CTSG and NYSG, respectively,) and the U.S. Environmental Protection Agency (EPA) through the Long Island Sound Study (LISS), will attempt to answer questions critical to advancing restoration of the estuary and its watershed. All the awards are supplemented with matching funds of at least 50 percent of the grants, extending the value of the research package to more than $4.2 million.
All the projects will span two years, with work slated to begin this spring.
The eight research projects are the latest to be awarded through the Long Island Sound Study Research Grant Program, run by NYSG and CTSG since 2008. Including the new awards, the program has funded ecological research in more than 30 areas. It represents the largest research investment into the Sound, designated an estuary of national significance and one of the most valuable natural resources for both states.
The projects are:
- “Can Watershed Land Use Legacies Inform Nitrogen Management?” (University of Connecticut, Ashley Helton, Chester Arnold, Emily Wilson, David Bjerklie; University of New Hampshire, Wilfred Wollheim; CT DEEP, Mary Becker, Chris Bellucci; Footprints in the Water, Paul Stacey: $487,391). This project will examine the impact of historical land use practices in managing nitrogen.
- “Evaluating Thin Layer Placement in Long Island Sound Marshes Using a Multi-Scale Approach” (University of Connecticut, Beth Lawrence, Ashley Helton, Chris Elphick; CT DEEP, Min Huang: $470,969.) In this project, different types of sediment for effective marsh rebuilding will be assessed.
- “Can They Get Out? Assessing the Effects of Low Streamflow on Juvenile River Herring” (University of Connecticut, Eric Schultz, James Knighton, Cary Chadwick: $231,013.) Researchers in this project will identify barriers to the outmigration of juvenile alewife, a keystone species in the Sound’s food chain.
- “Establishing Robust Bioindicators of Microplastics in Long Island Sound: Implications for Reliable Estimates of Concentration, Distribution and Impacts” (University of Connecticut, J. Evan Ward and Sandra Shumway, $301,150.) One of two projects examining how different types of marine life can contribute to efforts to quantify and remove pollutants, this research will examine the use of slipper snails, tunicates and oysters as bioindicators of the concentrations and impacts of microplastics.
- “Quantifying the Ability of Seaweed Aquaculture in Long Island Sound to Remove Nitrogen, Combat Ocean Acidification, Improve Water Quality and Benefit Bivalves” (Stony Brook University, Christopher Gobler, Michael Doall; GreenWave, Kendall Barbery: $238,933.) In this project, the ability of cultured seaweed and shellfish to remove nitrogen, combat ocean acidification, improve water quality and benefit aquaculture will be measured.
- “Constraining Models of Metabolism and Ventilation of Bottom Water in Long Island Sound Using Oxygen Isotopes” (University of Connecticut, Craig Tobias, James O’Donnell: $694,386.) One of three projects focusing on conditions in water and sediment chemistry, this work will examine factors that influence recovery from hypoxia (low oxygen).
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An eelgrass bed off of Fishers Island, N.Y. Eelgrass bed restoration will be the focus of one of the eight research projects. Photo: CCE Suffolk County Marine Program. “Improving Eelgrass Restoration Success by Manipulating the Sediment Iron Cycle” (University of Connecticut, Craig Tobias and Jamie Vaudrey; and Cornell Cooperative Extension, Chris Pickerell: $323,404.) This project will evaluate methods to overcome sediment conditions that may impede eelgrass recovery.
- “Alkalinity of Long Island Sound Embayments” (University of Connecticut, Penny Vlahos and Michael Whitney; Save the Sound, Peter Linderoth; CT DEEP, Katie O’Brien-Clayton: $131,088.) This research will evaluate the vulnerability of embayments to changes in acidity that can be harmful to shellfish and other marine life.
Descriptions of all the Long Island Sound Study research projects, since the first grant cycle in 2000, are available in the research section of the LISS website.
In Their Words: Long Island Sound Study Research Projects
“EPA has a longstanding commitment to restoring and protecting Long Island Sound, one of the country’s most important estuaries. This funding will advance ecological research and play a critical role in improving water quality and reducing pollution, providing lasting results for the wildlife and wetlands in the Sound for years to come.” — acting EPA New England Regional Administrator Deb Szaro
“More than 10 percent of Americans live within 50 miles of the Long Island Sound’s shores, where issues like nitrogen pollution threaten water quality, marine life and coastal resiliency. These projects reflect EPA’s longstanding commitment to developing solutions to protect and restore the Sound to healthy waters, benefitting surrounding communities environmentally, economically and recreationally.” — EPA Region 2 acting Regional Administrator Walter Mugan
“This research competition resulted in an interesting diversity of projects. These include novel approaches to understanding and managing Long Island Sound and reaching the goals of increased water quality that support productive ecosystems for the benefit of wildlife and humans. In my opinion, it is a very smart investment for long-term benefits.” — Connecticut Sea Grant Director Sylvain De Guise

“New York Sea Grant is pleased to continue our longstanding partnership with Connecticut Sea Grant and the EPA through the LISS competitive research program. The Long Island Sound is an essential and beloved natural asset to the citizens of NY and CT and supports critical environment and human ecosystem services. The eight diverse and innovative research initiatives that were awarded funding will provide critical knowledge needed to ensure the health of the Sound today and into the future.” — New York Sea Grant Director Rebecca Shuford
“This grant cycle represents the largest amount of funding ever competed and resulted in a very competitive and diverse group of projects. This research effort is even more significant because it is tightly linked to management applications so will go far to improve our understanding of the Sound and help managers in their work to restore and improve the Sound’s waters and biota.” — CTSG Research Coordinator Syma Ebbin
“This cohort of projects continues the successful long-term collaboration between CTSG, NYSG and EPA LISS that continues to expand the knowledge base that is invaluable to aiding the management of Long Island Sound. The results of this research effort will benefit the LIS environment and its communities.” — NYSG Research Coordinator Lane Smith
For more information, contact CT Sea Grant Communications Coordinator Judy Benson.